This circuit looks a bit like a reference current generator I sometimes use:
It also works with bipolar transistors so ignore the fact that this circuit uses MOSFETs. Also imagine that R is your sense resistor and the load resistor.
This circuit works by having a 1 : 1 current mirror (M3 and M4) which makes the left and right currents equal. But M1, M2 and R also make a mirror but it is not 1 : 1, it is actually non-linear.
Having these two work together means a solution must satisfy both mirrors.
The wanted solution is where left and right currents are equal but there is also the solution where all currents are zero.
So this circuit needs a start-up circuit to detect that and pull it to the wanted (1:1) solution.
I think that is where your R3 and R4 come in. These force a current through Rsense, R2 and T2 so that the circuit can start up properly.
Once started the current through R3 and R4 will be small, much smaller than the nominal current the circuit is designed to run at. That explains the relatively high value of R3 and R4.
I would have drawn R3 and R4 not up but down from the base of T2 so that it is clear they conduct a current to ground.