Basic idea
A capacitor is neither an open circuit nor a short connection; it is a "duplicating voltage source" (a "voltage clone"). Imagine the simplest capacitive circuit - a capacitor connected to a DC voltage source. The capacitor is charged to the source voltage and no current flows in the circuit because two sources of equal but opposite voltage are connected in a loop.
Operation
Emulated capacitor: We can actually replace the charged capacitor with a voltage source Vc and thus conveniently examine it through an DC ammeter and the CircuitLab DC Live simulation.
DC input voltage
If you start to slowly change the source voltage, the "capacitor" will also change its voltage; their difference will remain equal to zero and no current will flow. We can mimic this by setting the same voltages to both Vin and Vc:
Vin = 1 V; Vc = 1 V -> Ic = 0
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Vin = 2 V; Vc = 2 V -> Ic = 0
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Vin = 3 V; Vc = 3 V -> Ic = 0
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So the source has the illusion that there is nothing connected to it, but it is, and it is a voltage source. There is also an ammeter connected with almost zero resistance, but the source "sees" an infinitely high resistance as if by some magical means the ammeter resistance is increased many times over. This trick is known in circuitry as bootstrapping.
Slowly changing DC input voltage
Conceptual circuit: We can automate this experiment by replacing the input voltage source and the emulated capacitor by two equal low-frequency AC voltage sources.
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As above, their difference remains equal to zero and again no current flows.
Practical circuit: Let's finally explore a true capacitive circuit. Since the capacitance is small enough, the capacitor quickly charges...
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... and its voltage follows the input voltage; the current is low.
AC input voltage
If you start rapidly changing the input voltage, the "capacitor" will not be able to change its voltage Vc, and the current will change significantly. To limit the current to a reasonable value and to "cheat" the simulator, I have set a low resistance (only 1 Ω) to the ammeter.
Vin = 2 V, Vc = 1 V, Ic = 1 A
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Vin = 3 V, Vc = 1 V, Ic = 2 A
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Vin = 4 V, Vc = 1 V, Ic = 3 A
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Thus the input source will have the illusion of being shorted, but in fact it is not.
Swept input voltage: To automate the experiment, let's sweep the input voltage around Vc...
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AC practical circuit: ... and finally explore the real capacitive circuit with an AC input voltage.
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As you can see, Vin varies only with +100 mV above and below 2 V...
... but the current significantly changes from -500 mA to 500 mA.
Generalization
Now we can imagine what happens in the more complex circuits with capacitors.
Capacitor acting as an open circuit
When the power supply is turned on, the capacitors are charged to the (quiescent) voltages between the respective nodes that would be without capacitors. There are no voltage differences, currents do not flow, as if the capacitors are disconnected; figuratively speaking, the circuit "softens". This open-circuit capacitor property is used when the operating (quiescent) point is set.
Capacitor acting as a short circuit
As the regulating element begins to vary its current, the voltages between the nodes begin to change. Currents begin to flow and the capacitors are "connected" to the circuit; figuratively speaking, the circuit "hardens". This short-circuit capacitor property is used when an input AC voltage (no matter with small or large amplitude) is applied.
Applications
Let's consider (the output part of) a typical circuit of an AC common-emitter amplifier stage with blocking capacitor in the emitter and coupling capacitor between the collector and load. In this conceptual circuit, I have emulated the collector-emitter (drain-source) part of the transistor with variable resistor Rce. Thus a voltage divider is formed by the three resistors Rc, Rce and Re.
Initial state
By adjusting Rce we set the desired quiescent collector voltage (10 V) and emitter voltage (5V).
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Connecting voltmeters
Then let's measure the voltages between the node pairs by connecting voltmeters in the place of the future "capacitors" (voltage sources). Remember their readings.
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Connecting "capacitors"
Then replace the voltmeters by voltage sources that emulate the capacitors. Adjust the source voltages equal to the voltmeter readings above ("charge the capacitors"). As you can see, nothing changes (hover over the circuit devices to see the voltages and currents by the help of the CircuitLab DC Live-Simulation feature).
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Decreasing Rce
Now decrease the resistance Rce of the regulating element ("transistor"). The divider's current increases so the emitter voltage across Re increases and becomes higher than the capacitor voltage Ve. The "capacitor" is "connected" and the divider passes a current through it.
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At the same time, the voltage drop across Rc increases. The voltage between the "transistor collector" and load RL decreases and becomes lower than the capacitor voltage Vc. The "capacitor" is "connected" and it passes a current through the divider and load RL, but since the "capacitance" is significant, the voltage across the "capacitor" almost does not change. So it transfers the "shifted" voltage decrease to the load, and the load voltage goes below zero.
Increasing Rce
Then increase Rce. The divider's current decreases so the emitter voltage Re decreases and becomes lower than the capacitor voltage Ve. The "capacitor" passes a current through Re but its voltage almost does not change.
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The voltage drop across Rc also decreases. The voltage between the "collector" and load RL increases and becomes higher than the capacitor voltage Vc. The divider passes a current through the capacitor and load RL but the voltage across the "capacitor" almost does not change. So it transfers the voltage increase and the load voltage goes above zero.
Thus the "collector voltage" variations around the 10 V quiescent voltage are "moved" to around ground.